Dr. Jörg Freyhof

IGB, Dept.4,
Biology and Ecology of Fishes 
12587 Berlin, Müggelseedamm 310

Fields of Interests:

Organisation of aquatic biodiversity

Tel: (030) 64 181 613
Fax: (030) 64 181 750

freyhof@igb-berlin.de

· “Long-term fluctuations of fish-community structures in the River Rhine basin.“

· “Diversity and perspectives of sustainable use of the fish fauna in the Central Mountain massifs of Vietnam.”

a) Factors structuring fish communities

Since 1992, ecological factors structuring the distribution, abundance and recruitment of fish species within a river drainage are studied in the premontane River Sieg, a main tributary of the River Rhine. The Sieg is an almost free flowing, small (33 m³/sec.) epipotamal river with strong floods, especially during winter. This river is inhabited by a rich fish fauna (41 spp.) including invasive species such as Aspius aspius and Vimba vimba. Our long term study is focused on the question how species specific recruitment and fish community structure are determined directly and indirectly by climate factors, taking “global warming” into account. In the River Sieg, recruitment strength, measured as juvenile fish abundance in summer, depends mostly on discharge in spring, favouring early spawners (e. g. Leuciscus leuciscus and Chondrostoma nasus) if water level is low. Suitable conditions for early spawning, predatory A. aspius seem to lead to a high predation pressure on juveniles of late spawning prey species (especially Alburnus alburnus). Water level fluctuations determine the habitat availability for juvenile fish: if water level is low, large amounts of very shallow lotic habitats are available. Late spawning species seem to be much less affected by high spring discharge in their year class strength compared to early spawners. The correlation between recruitment strength and adult fish abundance is weak in most species. This might be due to severe bottlenecks which determine the juvenile fish abundance, especially in winter. The abundance of most species seems to be limited by the amount of predator-poor refuges during winter floods. Winter habitat availability appears to be a major bottleneck for fishes smaller than 15 cm TL. Successful overwintering of some species is negatively correlated to year-class strength of perch Perca fluviatilis. Due to the fact that suitable winter habitats are more abundant in the lower reaches of the River Sieg, there are large upriver migrations in some species in spring. Furthermore, some species highly depend on winter habitats in the River Rhine. River Sieg is functioning as spawning river and source of juvenile fish for the main Rhine.

River Sieg at the village Herchen, a typical river of the barbel zone with excellent research opportunities.

b) Species pairs and sympatric speciation

The theory of sympatric speciation suggests that in a heterogeneous environment, individual fish can form different ecological niches. The specialization to one of these niches and the performance of different „life-history traits“ within a given environment should lead to disruptive evolution to solve the different ecological problems related to different niches. We study the ecological factors determining the origin and forcing the divergence of species pairs in Baltic ciscoes (Coregonus spp.) in lakes of Northern Germany. The occurrence of sympatric Baltic ciscoes having different spawning seasons or body sizes was already recognised by early authors in several post-glacial lakes. Our study supports the hypothesis of an independent, sympatric evolution of spring-spawning from autumn-spawning ciscoes. Two cases of sympatric speciation were found, involving two endemic species in Lakes Stechlin (Brandenburg) and Breiter Luzin (Mecklenburg-Vorpommern). Lake Breiter Luzin contains two ‘normal’-growing species, whereas in Lake Stechlin, a ‘normal’-growing and a dwarf species were found.

The species pair of Lake Stechlin: a, Coregonus albula; b, C. fontanae

Species pairs of ciscoes may be good models to study the mechanisms of sympatric speciation. Few studies have convincingly linked ecological factors and species formation to a point where key factors which are driving the divergence could be identified. The ecological processes leading to sympatric speciation in two pelagic species as ciscoes have not been uncovered so far. Generally, the distribution of spring spawning ciscoes is restricted to relatively deep and small lakes. However, only incomplete data are available on differential distribution patterns of species pairs within a lake. These data support that the spring spawning ciscoes inhabit greater water depths than the autumn spawning ones. Sympatric species pairs of ciscoes are known from about 14 Baltic lakes in Scandinavia and Russia. All these species pairs are presumed to originate from one or two invading species and might all be of the same age and sympatric origin. The morphology of at least one divergent species in the pairs should be closely related and adaptive to the individual lake environment. We search for general evolutionary patterns forming the ecology and morphology of species pairs as a reaction to particular lakes ecosystems. Ongoing research intends to test detailed hypotheses on life history differentiation and adaptive divergence. Actually, we conduct field research in Lake Stechlin Ingeborg Palm Helland and laboratory tests Jan Ohlberger and Georg Staaks to determine effects of abiotic (temperature, water depth) and biotic (food, competition, mating) factors to retrace the ecological forces which lead to sympatric speciation.

Position of Lake Stechlin

Together with Ingeborg Palm Helland and Thomas Mehner and covered by the German Science Foundation (DFG-Schwerpunkts Aquashift; DFG Me 1686/5-1) the ecology of Lake Stechlin ciscoes is intensively studied at the moment.

c) Ecological gradients and parapatric speciation

Specific „life history“ strategies may lead to the evolution of closely related species along ecological gradients. We study sculpins (Cottus spp.) from the lower River Rhine drainage in cooperation with Arne Nolte. Two species of Cottus occur in a parapatric pattern within the lower Rhine drainage.

Cottus perifretum, an invasive sculpin in the lower Rhine.

Apart from morphological and genetic differentiation, the two species strongly diverge in their potential to colonize subregions within an ecological continuum (pdf). They may be a good model to study the mechanisms of parapatric speciation along gradients. It should be possible to identify ecologically relevant traits upon which selection acts. While Cottus perifretum lives in warm, large rivers, Cottus rhenanus is restricted to cold headwaters. Yet, the two species form multiple extremely narrow hybrid zones in areas where small streams enter larger rivers. Both species hybridise freely but hybrids are confined to the hybrid zones. Pure species do not expand into the areas colonized by the other. The clear pattern at hybrid zones strongly suggests a key role of differential adaptation and external selection pressures in maintaining the genetic divergence. Ecological data indicate a strong differentiation of both species in life-history patterns. Cottus perifretum is more r-strategic, which might allow this species to colonize the stochastic and fecund potamal habitats with high predation pressure whereas predation might to be too high for slowly reproducing Cottus rhenanus in this habitat. In streams Cottus rhenanus might be the better competitor. Current research tests the hypothesis that life history differentiation lead to adaptive divergence. We conduct transplantation experiments in natural habitats and laboratory tests to determine effects of abiotic (temperature) and biotic (predation, food, physiology) factors.

Barbatula barbatula inhabits a long environmental gradient from small highland streams to medium sized lowland rivers.

Stone loaches (Barbatula) might be an ideal model system to study speciation along environmental gradients. This species is continuously distributed in a wide stream-river gradient and they show a very limited mobility. Preliminary studies show, that loaches from the lower and upper part of a river examined belong to one phylogeographic group. The differences in life history patterns of loaches from a stream and from a river population let us hope, that we could be able to find the interesting early stages of the environmental gradient speciation scenario.

In future, it will be studied, if life history differences integrate continuously or stepwise and we will search for the forcing ecological factors behind the differences (Climate, Predators).

d) Diversity and Conservation of Freshwater fishes

Freshwater fishes of Europe

Biodiversity management and conservation is one of the few global political goals that have been agreed upon at the highest possible political level. The loss of biodiversity, i. e. genetic and species diversity represents a loss to present and future generations. Therefore, the protection of biodiversity and the buffering of negative effects of increasing economic development must be given high priority within society. As species are bearers of genetic diversity and building blocks of ecosystems, information on their conservation status and distribution is fundamental to making competent decisions about protection of biodiversity from the local to global scales.

Surprisingly, the fish fauna of Europe is still incompletely known. In Europe, 547 native and 33 alien species are recorded in 2007. Since the year 2000, 57 species (10% of Europe’s fauna) have been described as new. Still many undescribed species are known or suspected to exist in all parts of Europe, including Germany and the number of European species might easily rise up to 600 or more. Within the framework of IUCN’s global fish red listing, we have completed the assessment the conservation status of all freshwater fishes endemic to the Mediterranean (see link) and all freshwater fishes native to Europe. This is an important step in building up a database for freshwater fish conservation for the EU action plan: Halting biodiversity loss by 2010 (see link), the EU Water Framework Directive, Natura 2000 and many national and regional activities. The IUCN Red List helps to find conservation priorities for European freshwater fishes. Creating widespread awareness of the values of threatened fishes and helping action plans to be started is one of the aims of the IUCN/SSC Freshwater specialist group.

European fishes in Turkey and Iran

The major gap in the evolutionary and biogeographic study of Europe’s freshwater fishes are now the fishes of Turkey and Iran. Biogeographically, both countries clearly belong to Europe having a very similar fauna and very similar problems in conservation. Many European fishes occur also in these countries and some species have the major part of their populations in Turkey or in Iran. Freshwater fishes of Turkey and Iran are poorly known but several of them seem to be at the very border of extinction. Since 2006, we cooperate with Müfit Özulug from Istanbul Üniversitesi and since 2007 with Ashghar Abdoli from Shahid Beheshti University to assess the biodiversity and the conservation status of Turkish and Iranian freshwater fishes.

Estinct: Coregonus oxyrinchus, the houting from England and the Rivers Rhine, Meuse and Scheldt.

German fishes

“Common wisdom” is that well-developed countries, such as Germany, are at the forefront of “the biodiversity issue”, setting examples for having their biota well documented and for enforcement of local management and conservation practices at the national level. Against this background, the recognition that aquatic biodiversity in Germany, even of key groups such as commercially important fish, is poorly known is highly alarming. Lack of knowledge of native diversity effectively prevents recognition of the magnitude of the problem that rapid environmental change and habitat loss enforce on aquatic biodiversity in Germany and elsewhere.

Until now, we can only estimate that the German ichthyofauna (check list) comprises some 106 freshwater species (including the anadromous and catadromous species), including 14 non-native species more or less established. Distribution data of German freshwater fishes are now available from the German digital fish species atlas (http://www.fischartenatlas.de). During the 19th and the 20th century, 12 species disappeared from Germany or became extinct. Three of our native species are globally extinct (Salvelinus profundus, Coregonus gutturosus, C. oxyrinchus). At least 12 species, all lacustrine, are endemic or almost endemic to German waters (pdf). Most likely, there are more endemic and more extinct species on the German territory. Especially in the subalpine lakes, human impact may have resulted in a high number of extinctions of endemic fish species. This might be the biggest post-glacial extinction crisis of vertebrate species in Central Europe.

Survived: Salvelinus evasus, the only real deep-water charr from the alpine lakes.

Actually, we try to close the gaps of knowledge in national biodiversity and try to reconstruct the biogeographic mechanisms, which lead to the actual fish distribution in Germany. Since most of our native species had their glacial refuges in Eastern Europe, we cooperate with the Zoological Institute St. Petersburg to investigate the biogeography and conservation status of freshwater fishes in the Baltic and Black Sea basins, the most diverse basins in Europe. Because molecular methods allow analysing biogeographic patterns, we actually cooperate with Arne Ludwig, Jörg Bohlen & Vendula Slechtova.

Alburnus mento, a shemaya endemic to the subalpine lakes of Austria and Germany.

to head of document

„Ökologie von Flussauen am Beispiel des Rheins“ (Seminar im WS 1995/96)

„Standardisierte Methoden zur Erfassung von Tierartengruppen für die Planung“ (Seminar im WS 1995/96; 1996/97)

„Limnologie von Fließgewässern“ (Seminar im WS 1996/97)

„Erhebung, statistische Auswertung und Präsentation feldbiologischer Daten“ (Blockkurs im WS 1996/97; 1997/1998)

„Diversität und Ökologie von Fischen“ (Seminar im WS 1997/98)

Humboldt-Universität Berlin, Mathematisch-Naturwissenschaftliche Fakultät

„Ökologie eines Flussystems am Beispiel der Elbe“ (Seminar im SS 2003)

„Speziation der Fische - Artbildungsprozesse unter Wasser“ (Seminar im WS 2003/04; 2004/05)

„Life in the cold - Wirbeltiere im Winter” (Exkursionen im WS 2003/2004; 2004/2005)

„Versuchsplanung und Auswertung freilandbiologischer Daten“ (Blockkurs im WS 2003/04; 2004/2005)

„Ökologie und Diversität der Fische“ (VL im WS 2004/05)

„Ökologie und Diversität der Fische“ (VL im WS 2004/05)

Universität Potsdam, Mathematisch-Naturwissenschaftliche Fakultät, Institut für Biochemie und Biologie

„Ökologie und Diversität der Fische“ (VL im SS 2004 with mit Franz Hölker)

Students playing with Coregonus at Lake Stechlin, April 2005

to head of document

Publikations

Handbook of European Freshwater fishes

Kottelat, M. & Freyhof, J. 2007. Handbook of European freshwater fishes. Kottelat, Cornol and Freyhof, Berlin, xiv + 646 pp.

The Handbook of European Freshwater Fishes provides the most up to date taxonomic review, identification key and ecology and distribution information for all 546 native (and 33 introduced) freshwater fish species in Europe.

The book contains:

– a key to genera and species
– diagnoses for all species
– methods for identification
– habitat, biology and ecology
– native, extirpated and introduced distribution information and maps
– conservation status, (using IUCN Red List Categories and Criteria)
– up-to-date taxonomy and nomenclature, with modern methods and concepts
– more than 870 bibliographic references
– more than 900 illustrations
– colour photographs for nearly every species

It is „THE“ book about the diversity and biology of European freshwater fishes.

Please click here to download the order form

Please click here to download the order form

Please click here to download the order form

Journal Publications

1. Freyhof, J. 1995. Zum Vorkommen der Gemeinen Keiljungfer Gomphus vulgatissimus (L.) (Odonata, Gomphidae) an der Sieg/NRW. Decheniana 148, 121-123.

2. Weiler, W., Freyhof, J. & M. Schulz 1995. Zum Vorkommen von Branchipus schaefferi Fischer (Crustacea, Anostraca) in der Wahner Heide. Decheniana, 148, 120.

3. Freyhof, J. 1995. Feeding traces as an indicator of habitat choice in nase Chondrostoma nasus (L.). Folia Zoologica 44 (Suppl.1), 57-61.

4. Freyhof, J. 1996. Veränderungen der Gattungszuordnung bei zwei mitteleuropäischen Süßwasserfischen. Fischökologie 9, 60-61.

5. Freyhof, J. 1996. Die Laichpopulation des Flußneunauges Lampetra fluviatilis L. in der unteren Sieg. Fischökologie 9, 80-81.

6. Freyhof, J. 1996. Distribution of YOY-Barbel Barbus barbus (L.) in the River Sieg/Germany. - In: Kirchhofer, A. & Hefti, D. Edit. Conservation of endangered freshwater fish in Europe. Advances in Life Sciences, Birkhäuser, 259-267.

7. Persat, H., Beaudou, D. & J. Freyhof 1996. The sculpin of the Lez spring (South France), Cottus petiti (Bacescu and Bacescu-Mester, 1964), one of the most threatened fish species in Europe. - In: Kirchhofer, A. & Hefti, D. Edit. Conservation of endangered freshwater fish in Europe. Advances in Life Sciences, Birkhäuser, 321-328.

8. Freyhof, J. 1997. Zur Fortpflanzungsbiologie des Barbengründlings. - DATZ 50: 513-516.

9. Freyhof, J. 1997. Age-related longitudinal distribution of nase Chondrostoma nasus in the River Sieg, Germany. Folia Zoologica 46/Suppl. 1, 89-96

10. Freyhof, J. 1997. Remarks on the status of Chondrostoma nasus (L.) in the system of the river Rhine. Folia Zoologica 46/Suppl. 1, 61-66

11. Steinmann, I. & J. Freyhof 1997. Body conditions of Chondrostoma nasus (L.) during first winter - Folia Zoologica 46/Suppl. 1, 117-122

12. Freyhof, J. & Steinmann, I. 1998. Rhithropanopeus harrisii (Gould 1841) und Palaemon longirostris Edwards im Rhein bei Köln (Crustacea, Decapoda). Lauterbornia 32, 25-26

13. Freyhof, J., Steinmann, I. & Krause T. 1998. Weitere Funde von Gomphus flavipes (Charpentier) im Rhein (Anisoptera: Gomphidae). Libellula 17, 247-252

14. Freyhof, J. 1998. Exuvien- und Larvenfunde der Asiatischen Keiljungfer Gomphus flavipes (Charpentier) im Rhein von Nordrhein-Westfalen. LÖBF-Mitteilungen, 4, 4-4.

15. Bischoff, A., Freyhof, J. & Staas, S. 1998. Nachweise des Zobels Abramis sapa (Pallas, 1811), (Teleostei: Cyprinidae) im Rhein. Lauterbornia, 33, 5-8

16. Freyhof, J. 1998. Die Fische und Neunaugen der Sieg in den Grenzen von Nordrhein-Westfalen. Decheniana, 151, 183-194

17. Steinmann, I.; Freyhof, J. & Busse, K. 1998. Der Silberling „Leuciscus dolabratus Hollandre, 1838„ - ein wenig bekannter Cyprinidenhybride. Verh. Ges. Ichthyol. 1, 219-223.

18. Freyhof, J., Staas, S. & Steinmann, I. 1998. Erste Nachweise des Weißflossengründlings Gobio albipinnatus Lukasch, 1933 im Rhein (Telostei, Cyprinidae). LÖBF-Mitteilungen, 3, 75-77.

19. Freyhof, J. 1998. First record of Pomatoschistus canestrinii (Ninni, 1883) in Lake Trasimeno. Riv. Idrobiol., 37: 107-108.

20. Freyhof, J. 1998. Strukturierende Faktoren für die Fischgemeinschaft der Sieg. Cuvillier Verlag Göttingen, 164 pp.

21. Freyhof, J. 1998. Zur Unterscheidung der Ammocoeten des Meerneunauges Petromyzon marinus L. und Flußneunauges Lampetra fluviatilis (L.) im Feld. Österr. Fischerei 51, 167-168

22. Freyhof, J. 1998. Wissenschaftliche Namen zweier europäischer Süßwasserfische. Fischökologie Aktuell 11, 2-7

23. Freyhof, J. & Steinmann, I. 1998. Winterliche Verteilungsmuster von Fischen in der Sieg. LÖBF-Mitteilungen, 2, 56-58

24. Freyhof, J. 1998. Riffle spawning white bream Abramis bjoerkna in a regulated river - a case of unsuitable habitat choice?. Ital. J. Zool. 65 (Suppl.), 441-444

25. Bischoff, A. & Freyhof, J. 1999. Seasonal shifts in day-time resource use of 0+ barbel Barbus barbus. Environmental Biology of Fishes 56, 199-212.

26. Busse, K. , Freyhof, J. & Zarske, A. 1999. Serum-electrophoretic patterns of various cyprinid hybrids from the rivers Elbe and Sieg (Teleostei: Cyprinidae). Faun. Abh. Mus. Tierk. Dresden 21, 265-272.

27. Freyhof, J. & Weibel, U. 1999. Vorschlag einer Neufassung der Rote Liste der bestandesgefährdeten Neunaugen und Fische (Petromyzontiformes; Teleostei) in Rheinland-Pfalz sowie ein Verzeichnis aller vorkommender Arten. Fauna und Flora in Rheinland-Pfalz 9: 195-206.

28. Freyhof, J. 1999. Records of Vimba vimba (L.) (Teleostei: Cyprinidae) in the River Rhine and its tributaries. Folia Zoologica 48: 84-89.

29. Wolter, C., Bischoff, A. & Freyhof, J. 2000. Vorkommen und Verbreitung des Weißflossengründlings Gobio albipinnatus Lukatsch 1933 in der unteren Oder. Gewässerökologie Norddeutschlands, 4: 137-138.

30. Englbrecht, C., Freyhof, J., Nolte, A., Rassmann, K., Schliewen, U. & Tautz, D. 2000. Phylogeography of the bullhead Cottus gobio (Pisces: Teleostei: Cottidae) suggests a pre-Pleistocene origin of the major Central European populations. Molecular Ecology 9: 709-722.

31. Freyhof, J., Scholten, M., Bischoff, A., Wanzenböck, J., Staas, S. & Wolter, C., 2000. Extensions to the known range of the whitefin gudgeon Gobio albipinnatus Lukasch, 1933, in Europe and biogeographical implications. J. Fish Biol. 57. 1339-1342.

32. Freyhof, J. A. & Serov, D. V. 2000. Cobitis laoensis (Sauvage, 1878) from Vietnam with notes on the southern distribution limits of Indochinese Cobitis. Folia Zoologica 49 (Suppl. 1), 205-214.

33. Freyhof, J., Ráb, P., & Bohlen, J. 2000. The valid names of some European species of the genus Cobitis (Teleostei, Cobitidae). Folia Zoologica 49 (Suppl. 1), 3-7.

34. Freyhof, J. & Serov, D. V. 2000. Review of the genus Sewellia from Vietnam with descriptions of two new species (Cypriniformes: Balitoridae). Ichthyological Exploration of Freshwaters 11, 217-240.

35. Freyhof, J., Serov, D. V. & Nga, N. T. 2000. A preliminary checklist of the freshwater fishes of the River Dong Nai, Southern Vietnam. Bonner Zoologische Beiträge 49: 93-99.

36. Freyhof, J. & Serov, D. V. 2001. Nemacheiline loaches from Central Vietnam with descriptions of a new genus and 14 new species (Cypriniformes: Balitoridae). Ichthyological Exploration of Freshwaters 12, 133-191.

37. Freyhof, J. & Herder, F. 2001. Tanichthys micagemmae, a new miniature cyprinid fish from Central Vietnam (Cypriniformes: Cyprinidae). Ichthyological Exploration of Freshwaters 12, 215-220.

38. Ng, H. H. & Freyhof, J. 2001. A review of the catfish genus Pterocryptis (Siluridae) in Vietnam, with the description of two new species. J. Fish Biol. 59: 624-644.

39. Ng, H. H. & Freyhof, J. 2001. Oreoglanis infulatus, a new species of glyptosternine catfish (Siluriformes: Sisoridae) from central Vietnam. J. Fish Biol. 59, 1164-1169.

40. Vences, M., Freyhof, J., Sonnenberg, R., Kosuch, J. & Veith, M. 2001. Reconciling fossils and molecules? Possible Cenozoic divergence of cichlid fishes, and the biogeography of Madagascar. Journal of Biogeography 28, 1091-1100.

41. Freyhof, J. 2002. Freshwater fish diversity in Germany, threats and species extinction, pp. 3-22. In: Collares-Pereira, M. J., Coelho, M. M. & Cowx, I. G. Freshwater fish conservation: options for the future, Blackwell Publishers 462 pp.

42. Bohlen, J., Rab, P., Slechtova, V., Rabova, M., Ritterbusch, D., & Freyhof, J. 2002. Hybridogenous biotypes in spined loaches (genus Cobitis) in Germany with implications for conservation, pp. 311-321. In: Collares-Pereira, M. J., Coelho, M. M. & Cowx, I. G. Freshwater fish conservation: options for the future, Blackwell Publishers, 462 pp.

43. Freyhof, J. & Herder, F. 2002 Records of genus Hemimyzon in Vietnam, with the description of a new species (Cypriniformes: Balitoridae). Ichthyological Exploration of Freshwaters 13, 53-58.

44. Freyhof, J. & Herder, F. 2002. Review of the paradise fishes of the genus Macropodus in Vietnam, with description of two new species from Vietnam and southern China (Perciformes: Osphronemidae). Ichthyological Exploration of Freshwaters 13, 147-167.

45. Sindilariu, P. D., Bacalbasa-Dobrovici, N., Freyhof, J., Wolter, C., 2002. The juvenile fish community of the lower Danube and the Danube Delta. International Association for Danube Research 34: 517-526.

46. Dönni, W. & Freyhof, J. 2002. Einwanderungen von Fischarten in die Schweiz, Rheineinzugsgebiet. Mitteilungen zur Fischerei 72, Bundesamt für Umwelt, Wald und Landschaft (BUWAL), 89 pp.

47. Sindilariu, P. D. & Freyhof, J. 2003. Food overlap of benthic fishes in the Danube Delta, with special respect to two invasive gobiids (Teleostei: Gobiidae, Percidae, Cyprinidae). Lauterbornia 46: 149-157.

48. Freyhof, J. 2003. Immigration and potential impacts of invasive freshwater fishes in Germany. Berichte des IGB 17: 51-58.

49. Freyhof, J. 2003. Sewellia albisuera, a new balitorid loach from Central Vietnam (Cypriniformes: Balitoridae). Ichthyological Exploration of Freshwaters 14: 225-230.

50. Schulz, M. & Freyhof, J. 2003. Coregonus fontanae, a new spring-spawning cisco from Lake Stechlin, northern Germany (Salmoniformes: Coregonidae). Ichthyological Exploration of Freshwaters 14: 209-214.

51. Ng, H. H. & Freyhof, J. 2003. Akysis clavulus, a new species of catfish (Teleostei: Akysidae) from Central Vietnam. Ichthyological Exploration of Freshwaters 14: 311-316.

52. Bohlen, J., Šlechtová, V., Šanda, R., Kalous, L., Freyhof, J. Vukic, J. & Mrdak, D. 2003. Cobitis ohridana and Barbatula zetensis in the River Moraca basin, Montenegro: Distribution, habitat, population structure and conservation needs. Folia biologica (Krakow), 51 Suppl.: 147-153.

53. Wolter, C. & Freyhof, J. 2004. Diel fish movements in a temperate large lowland river. Journal of Fish Biology 64: 632-642.

54. Naseka, A. M. & Freyhof, J. 2004. Romanogobio parvus, a new gudgeon from River Kuban, southern Russia (Cyprinidae, Gobioninae). Ichthyological Exploration of Freshwaters 15: 17-23.

55. Bogutskaya, N. G., Movchan, Y. V., Freyhof, J. 2004. On Barbus waleckii Rolik, 1970 (Cyprinidae) in Ukraine with brief remarks on species from the genus Barbus Cuvier, 1816, distributed in the Dniestr and Vistula drainages. Vestnik Zoologii, Kiev, 38, 87-92.

56. Yahyaoui, A., Freyhof, J. 6 Steinmann, I. 2004. Diversite ichthyologique et biologie d’Anguilla anguilla (L., 1758) (Actinopterygii, Anguillidae) in the middle Rhine. Zoologica Baetica 15, 39-60.

57. Šlechtová, V., Bohlen, J., Freyhof, J., Persat, H., & Delmastro, G. 2004. The Alps as barrier to dispersal in cold-adapted freshwater fishes? Phylogeographic history and taxonomic state of the bullhead in the Adriatic freshwater drainage. Molecular Phylogenetics and Evolution 33: 225-239. (4,213)

58. Freyhof, J. & Herder, F. 2004. Comments on the proposed conservation of the specific name of M. concolor Ahl, 1937 (Osteichthyes, Perciformes). Bulletin of Zoological Nomenclature 61 (3): 173-174.

59. Özulug, M., Meriç N. & Freyhof, J. 2004. The distribution of Carassius gibelio (Bloch, 1782) (Teleostei: Cyprinidae) in Thrace (Turkey). Zoology in the Middle East 31: 63-66.

60. Freyhof, J. & Brunken, H. 2004. Erste Einschätzung der Verantwortlichkeit Deutschlands für die Erhaltung von Fischarten und Neunaugen des Süßwassers. Naturschutz und Biologische Vielfalt 8: 133-144.

61. Freyhof, J. & Korte, E., 2005. The first record of Misgurnus anguillicaudatus in Germany. Journal of Fish Biology 66 (2): 568-571.

62. Freyhof, J. & Kottelat, M. 2005. Salvelinus evasus sp. n., a charr from deep waters of Lake Ammersee, southern Germany (Teleostei: Salmonidae), with comments on two extinct species. Revue Suisse de Zoologie 112, 1: 253-269. (0,284)

63. Bohlen, Freyhof, J. & Wolter, C. 2005. First records of Cobitis elongatoides and Sabanejewia baltica (Cobitidae) for Germany. Cybium 29(1): 103-104. (0,312)

64. Freyhof, J., Kottelat, M. & Nolte, A. 2005. Taxonomic diversity of European Cottus with description of eight new species (Teleostei: Cottidae). Ichthyological Exploration of Freshwaters 16: 107-172.

65. Freyhof, J. 2005. Redescription of Coregonus bavaricus Hofer, 1909, an endemic species to Lake Ammersee, Germany (Salmoniformes: Coregonidae). Cybium 29: 179-183. (0,312)

66. Herder, F. & Freyhof, J. 2005. Comment on the proposed conservation of the specific name of Macropodus concolor Ahl, 1937 (Osteichthyes, Osphronemidae). Bulletin of Zoological Nomenclature 62: 87-89.

67. Freyhof, J. & Schöter, C. 2005. The Houting, Coregonus oxyrinchus (L.), a globally extinct species from the North Sea basin (Salmoniformes: Coregonidae). Journal of Fish Biology 67: 713-729.

68. Copp, G.H., Bianco, P.G., Bogutskaya, N.G., Erős, T., Falka, I., Ferreira, M.T., Fox, M.G., Freyhof, J., Gozlan, R.E., Grabowska, J., Kováč, V., Moreno-Amich, R., Naseka, A.M., Peňáz, M., Povž, M., Przybylski, M., Robillard, M., Russell, I.C., Stakėnas, S., Šumer, S., Vila-Gispert, A. & Wieser, C. 2005. To be, or not to be, a non-native freshwater fish. Journal of Applied Ichthyology 21: 242-262.

69. Freyhof, J., Lieckfeldt, D., Pitra, C. & Ludwig A. 2005. Molecules and morphology: evidence for introgression of mitochondrial DNA in Dalmatian cyprinids. Molecular Phylogenetics and Evolution 37: 347-354.

70. Nolte, A., Freyhof, J., Stemshorn, K. & Tautz, D. 2005. An invasive lineage of sculpins, Cottus sp. (Pisces, Teleostei) in the Rhine with new habitat adaptations has originated by hybridization between old phylogeographic groups. Proceedings of the Royal Society, London, 272: 2379 - 2387

71. Ng, H. H. & Freyhof, J. 2005. Pseudomystus sobrinus a new catfish (Teleostei: Bagridae) from central Vietnam. Copeia, 744-749.

72. Kottelat, M., Bogutskaya, N. G. & Freyhof, J. 2005. On the migratory Black Sea lamprey and the nomenclature of the ludoga, Peipus and ripus whitefishes (Teleostei: Petromyzonidae and Coregonidae). Zootaxonomica Rossica, 181-186.

73. Freyhof, J. & Naseka, A. M. 2005. Gobio delyamurei, a new gudgeon from Crimean Peninsular (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters 16, 331-338.

74. Wolter, C. & Freyhof, J. 2005. Die Fischbesiedlung des Oder-Einzugsgebietes, Nationalpark-Jahrbuch Unteres Odertal, Nationalparkstiftung, Unteres Odertal, Schwedt, 2005: 37-63.

75. Naseka A. M., Spodareva V. V., Freyhof J., Bogutskaya N. G., Poznjak V. G. 2005. New data to species composition and distribution of gudgeons (Gobioninae, Cyprinidae, Teleostei) in River Kuban. Folia Zoologica 54, Suppl. 50-55.

76. Freyhof, J., Lieckfeldt, D., Bogutskaya, N. G. & Ludwig A. 2006. Phylogenetic position of the Dalmatian genus Phoxinellus and description of Delminichthys, new genus, (Teleostei: Cyprinidae). Molecular Phylogenetics and Evolution 38: 416-425.

77. Lieckfeldt, D., Freyhof, J. & Ludwig, A. 2006. Detection, characterization and utility of a new highly variable noncoding nuclear region in several species of cyprinid fishes (Cyprinidae). European Journal of Wildlife Research 52: 63-65

78. Yang, J., He, S., Freyhof, J., Witte, K.-E. & Liu, H. 2006. Phylogeny of the Gobioninae inferred from complete mitochondrial cytochrome b gene sequences (Teleostei: Cyprinidae). Hydrobiologia 553: 255-266.

79. Schulz, M., Freyhof, J., Saint-Laurent, R., Østbye, K., Mehner, T. Bernatchez, L. 2006. Evidence for independent origin of two spring-spawning ciscoes in Germany (Salmoniformes: Coregonidae). Journal of Fish Biology 68, Suppl. A: 119-135.

80. Sindilariu, P. D., Freyhof, J. & Wolter, C. 2006. Habitat use of juvenile fish in the lower Danube and the Danube Delta: implications for ecotone connectivity. Hydrobiologia, 571: 51-61.

81. Herder, F. & Freyhof, J. 2006. Niche differentiation of fishes in a medium-sized tropical stream. Journal of Fish Biology 68: 571-589.

82. Nolte, A. W., Freyhof, J., & Tautz, D. 2006. When invaders meet locally adapted types: rapid moulding of hybrid zones between two species of sculpins (Cottus, Pisces). Molecular Ecology15. 1983-1989..

83. Šlechtová, V., Bohlen, J., Freyhof, J. & Rab, P. 2006. Molecular phylogeny of the Southeast Asian freshwater fish family Botiidae (Teleostei: Cobitoidea) and the origin of polyploidisation in their evolution. Molecular Phylogenetics and Evolution 39: 529-541.

84. Bohlen, J., Slechtova, V., Bogutskaya, N. G. & Freyhof, J. 2006. Across Siberia and over Europe: Phylogenetic relationships of the freshwater fish genus Rhodeus in Europe and the phylogenetic position of R. sericeus from the River Amur. Molecular Phylogenetics and Evolution, 40: 856-865.

85. Freyhof, J. & Huckstorf, V. 2006. Conservation and management of aquatic genetic resources: a critical checklist of German freshwater fishes. Berichte des IGB 23: 113-126.

86. Freyhof; J. & Özulug, M. 2006. Pseudophoxinus ninae, a new species from Central Anatolia, Turkey (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 17: 255-259.

87. Stelbrink, B. & Freyhof, J. 2006. Reduction of scales and head canals in Pomatoschistus canestrinii (Ninni, 1883) (Teleostei, Gobiidae). Verhandlungen der Gesellschaft für Ichthyologie, 5: 71-77.

88. Helland, I. P., Freyhof, J., Kasprzak, P. & Mehner, T. 2007. Temperature sensitivity of vertical distributions of zooplankton and planktivorous fish in a stratified lake. Oecologia 151. 322-330.

89. Ng, H. H. & Freyhof, J. 2007. Pseudobagrus nubilosus, a new species of catfish from central Vietnam (Teleostei: Bagridae), with notes on the validities of Pelteobagrus and Pseudobagrus. Ichthyological Exploration of Freshwaters 18: 9-16.

90. Kottelat, M. & Freyhof, J. 2007. Pelasgus, a new genus name for the Balkan species of Pseudophoxinus (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 18: 103-108.

91. Kottelat, M. & Freyhof, J. 2007. Handbook of European freshwater fishes. Kottelat, Cornol and Freyhof, Berlin, xiv + 646 pp.

92. Helland, I. P., Brauns M., Freyhof, J. 2007. Depth distribution of abundant benthic invertebrates in Lake Stechlin. Berichte des IGB 24: 89-96.

93. Freyhof, J., & Kottelat, M. 2007. Alburnus vistonicus, a new shemaya from eastern Greece, with remarks on Chalcalburnus chalcoides macedonicus from Lake Volvi (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 18: 205-212.

94. Freyhof, J., & Kottelat, M. 2007. Review of the Alburnus mento species group with description of two new species (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 18: 213-225.

95. Özulug, M. & Freyhof, J. 2007. Rediagnosis of four shemajas from western Turkey and description of two new species (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 18: 233-246.

96. Freyhof, J. & Stelbrink, B. 2007. Cobitis illyrica, a new species of loach from Adriatic basin, Croatia (Teleostei: Cobitidae). Ichthyological Exploration of Freshwaters, 18: 269-275.

97. Mayden, R. L., Tang, K. L., Conway, K. W., Freyhof, J., Chamberlain, S., Haskins, M., Schneider, L., Sudkamp, M., Wood, R. M., Agnew, M., Bufalino, A., Sulaiman, Z., Miya, M., Saitoh, K. & He, S.-H. 2007 Phylogenetic relationship of Danio within the order Cypriniformes: A framework for comparative and evolutionary studies of a model species. Journal of Experimental Zoology 308B: 308B: 642–654.

98. Freyhof, J. & & Naseka, A. M. Proterorhinus tataricus, a new tubenose goby from Crimea, Ukraine (Teleostei: Gobiidae). Ichthyological Exploration of Freshwaters, 18: 325-334.

99. Özulug, M. & Freyhof; J. 2007. Alburnus demiri, a new species of bleak from Western Anatolia, Turkey (Teleostei: Cyprinidae). Ichthyological Exploration of Freshwaters, 18: 307-312.

100. Helland, I. P., Harrod, C., Freyhof, J. & T. Mehner. 2008. Coexistence of a pair of pelagic planktivorous coregonid fish. Evolutionary Ecology Research,

101. Tang, Q, Freyhof, J, Xiong, B, & Liu, H. 2008. Multiple invasions of Europe by East Asian cobitid loaches (Teleostei: Cobitidae). Hydrobiologia 605: 17-28.

102. Cano, J. M, Mäkinen, H. S, Leinonen, T, Freyhof, J. & Merilä, J. 2008. Extreme neutral genetic and morphological divergence supports Adriatic threespined stickleback (Gasterosteus aculeatus) populations as conservation units. Biological Conservation 141: 1055-1066.

103. Freyhof, J., Stelbrink, B., Özulug, M. & Economidis. P. S. 2008. The first record of Cobitis puncticulata from Europe with comments on its conservation status (Teleostei: Cobitidae). Folia Zoologica 57: 16–19.

104. Bohlen, J., Freyhof, J., Nolte, A. 2008. Sex ratio and body size in Cobitis elongatoides and Sabanejewia balcanica (Cypriniformes; Cobitidae) from a thermal spring. Folia Zoologica 57: 191–197.

105. Ng, H. H. & Freyhof, J. Two new species of Glyptothorax (Teleostei: Sisoridae) from Central Vietnam. Zootaxa, 1873: 11-25.

106. Lang, N. J., Roe, K. J., Renaud, C. B., Gill, H. S., Potter, I. C., Freyhof, J., Naseka, A., Cochran, P., Pérez, H. E., Habit, E. M., Kuhajda, B. R., Lyons, J., Neely, D. A., Reshtnikov, Y., Salnikov, V., Stoumboudi, M. T. & Mayden, R. L. Novel relationships among lampreys (Petromyzontiformes) revealed by a taxonomically comprehensive dataset. Transactions of the American Fisheries Society (in press)

107. Dümpelmann, C. & Freyhof, J. First record of the round goby Neogobius melanostomus (Pallas, 1814) in River Main, North Sea basin (Pisces: Gobiidae). Journal of Fish Biology (in press)

108. Freyhof, J. 2008. Rote Liste der im Süßwasser reproduzierenden Neunaugen und Fische (Cyclostomata & Pisces). Naturschutz und Biologische Vielfalt (in press)

Tagungsorganisation

„Freshwater fishes in tropical ecosystems: Threat and Protection” 9-12. March 1995 in Bonn, Zoologisches Forschungsinstitut und Museum Alexander Koenig.

„II International Symposium on the Biology of the Genus Chondrostoma Agassiz, 1835” 10.-13. October 1996 in Bonn, Zoologisches Forschungsinstitut und Museum Alexander Koenig.

„9th International Congress of European Ichthyologists (CEI)” 24-30. August 1997 in Trieste, (Member of International Steering Committee and Symposia Convenor, Part: Adaptive Ecology).

„11th International Congress of European Ichthyologists (ECI XI)” 6 - 10. September 2004 in Tallinn, Estonia, (Convenor of the Symposia 7. To be or not to be: freshwater fish conservation).

„12th International Congress of European Ichthyologists (ECI XII)” 9-13. September 2007 in Dubrovnik, Croatia, (Convenor of the Symposia. Conservation of freshwater fish).

Vorträge/Poster

„International Symposium Barbus III” 23.-26. June 1995 in Libnice/Cech Republic.

· Freyhof, J.: Habitat choice of Barbus barbus (L.) during first winter in the river Sieg, Germany. (oral)

„Jahrestagung der Deutschen Gesellschaft für Limnologie (DGL)“ 24.-29. September 1995 in Berlin/Germany.

· Bergfeld, T. & Freyhof, J.: Verteilungsmuster von 0+Jungfischen in Altwässern der Sieg, gefangen mittels eines aufschwimmenden Netzes. (poster)

· Scholten, M. & Freyhof, J.: Diurnale Aspekte der Nahrungsökologie larvaler Hasel und Nasen. (poster)

· Fielenbach, J. & Freyhof, J.: Zeitliche und räumliche Aktivitätsmuster der Nase Chondrostoma nasus (L.) in der Sieg. (poster)

· Weon, H.-Y. & Freyhof, J.: Drift und Makrozoobenthon in der Sieg. (poster)

· Freyhof, J.: Herbstliche Hochwasserschübe als bestimmende Faktoren für die Zusammensetzung und Verteilung von Jungfischgemeinschaften in der Sieg. (oral)

„Workshop zur Ökologie von 0+Fischen in großen Flüssen“ 30. September - 1. October 1995, Grietherbusch/Germany.

· Freyhof, J.: Herbstliche Hochwasserschübe als bestimmende Faktoren für die Zusammensetzung und Verteilung von Jungfischgemeinschaften in der Sieg. (oral)

· Fielenbach, J. & Freyhof, J.: Telemetrische Untersuchungen an Nasen in der Sieg - Erste Ergebnisse. (oral).

„Systematik, Ethologie und Ökologie der Fische“ 10.-12. February 1996, Salzburg/Austria.

· Freyhof, J: Saisonale Habitatverfügbarkeit für den Hasel Leuciscus leuciscus (L.) in der Sieg. (oral)

· Fielenbach, J. & Freyhof, J.: Telemetrische Untersuchungen an Nasen in der Sieg. (oral)

· Bischoff, A. & Freyhof, J.: Saisonale Verteilungsmuster innerhalb einer rheophilen Jungfischgemeinschaft in Rauschen der Sieg. (poster)

„Trophic interactions of age-0 fish and zooplankton in temperate waters” 5.-8. February 1996, Dresden/Germany.

· Bergfeld, T. & Freyhof, J.: Food of 0+roach and 0+bleak in backwaters of the river Sieg/Germany. (poster)

„II International Symposium on the Biology of the Genus Chondrostoma Agassiz, 1835” 10.-13. Oktober 1996, Bonn/Germany.

· Freyhof, J.: Status of Chondrostoma nasus (L.) in the system of the river Rhine. (oral)

· Freyhof, J.: The ecological profile of Chondrostoma nasus (L.) in the river Sieg. (oral)

· Fielenbach, J. & Freyhof, J.: Time and space utilisation modes by radiotagged Chondrostoma nasus in the river Sieg. (oral)

· Steinmann, I. & Freyhof, J.: Habitat choice and body conditions of Chondrostoma nasus during first winter. (poster)

„Global Biodiversity Research in Europe” International Senckenberg Conference, 9.-13. December 1996, Frankfurt/Germany.

· Freyhof, J.: Diversity and endangerment of cichlids (Teleostei, Cichlidae) in West-Africa. (oral)

„9. European Congress of Ichthyology ECI IX” 25.-29. August 1997, Trieste/Italy.

· Freyhof, J.: Riffle spawning white bream Abramis bjoerkna (L.) in a regulated river - a case of unsuitable habitat choice? (oral)

„1. Tagung der Gesellschaft für Ichthyologie (GfI)“ 9.-12. Oktober 1997, Düsseldorf/Germany.

· Steinmann, I.; Freyhof, J. & Busse, K.: Der Silberling „Cyprinus dolabrata Hollandre, 1838„ - ein wenig bekannter Cyprinidenhybride. (poster)

· Dalbeck, L.; Barenbroch, J. S. & Freyhof, J.: Abnutzungsmodus der Schlundzähne beim Döbel Leuciscus cephalus (L.). (poster)

· Dalbeck, L. & Freyhof, J.: Mechanismen der Inzestvermeidung beim Döbel Leuciscus cephalus (L.)? (poster)

· Freyhof, J. & Korte, E.: Das Meerneunauge Petromyzon marinus L. im Rhein und seinen Seitenflüssen. (oral)

„International Workshop: O+ fish as indicators of the ecological status of rivers” 16-20. 2. 1998 Wien /Austria.

· Freyhof, J.: Limited indicator value of the structure of 0+ fish assemblages. (oral)

„International Conference: Loaches of the genus Cobitis and related genera“ 31 May-3. June 1999, Brno/ Czech Republic.

· Freyhof, J.; Serov, D. V.: Southern distribution limits of Indochinese Cobitis in Vietnam. (poster)

„Eighth International Symposium of Regulated streams (EISORS)“ 17-21 July 2000, Toulouse/France.

· Freyhof, J.: Impacts of intensive ship traffic on the distribution and abundance of 0+ cyprinids. (oral)

„International Symposium: Freshwater fish conservation, options for the future“ 30. October - 4. November 2000, Albufera/ Portugal.

· Freyhof, J.: Freshwater fishes of Germany, theats and species extinction. (oral)

„10. European Congress of Ichthyology ECI X” 3. - 7. September 2001, Prague.

· Freyhof, J.: Riverine fishes in ecological research - is there a future? (plenary)

· Freyhof, J.: Fluctuations of the 0+ fish assemblage structure in the River Sieg, Germany. (oral)

· Herder, F. & Freyhof, J.: Biogeography of Central Vietnamese coastal river basins - one of the least known areas in Asia. (oral)

· Nolte, A., Freyhof, J. & Tautz, D.: Population genetics and morphological differentiation of bullhead Cottus gobio L. within the lower Rhine basin - persisting lineages as a result of postglacial speciation? (oral)

„94. Jahresversammlung der Deutschen Zoologischen Gesellschaft“ 4.-8. June 2001, Osnabrück.

· Nolte, A., Freyhof, J. & Tautz, D.: Population genetics and morphological differentiation of bullhead Cottus gobio L. within the lower Rhine basin - persisting lineages as a result of postglacial speciation? (oral)

“VIII International Symposium on the Biology and Management of Coregonid Fishes” 26.-29. August 2002, Rovaniemi, Finland.

· Schulz, M., Bernatchez, L., Freyhof, J., Garant, D., Rogers, S., Saint-Laurant, R., Ostbye, K. & Mehner, T.: Genetic evidence for parallel evolution of sympartic forms of Coregonus albula (L.) in two lakes of the Baltic lake area (Germany). (oral)

„IX Congress of European Society for Evolutionary Biology” 18. – 24. August 2003, Leeds, England.

· Nolte, A., Tautz, D. & Freyhof, J.: An apparent hybrid species of Cottus displays novel traits that faciliate colonization of new ecological subregions within the Rhine drainage. (oral)

· Nolte, A., Freyhof, J. & Tautz, D.: Hybrid zones of Cottus are fixed locally despite changes in population density. (oral)

„11. European Congress of Ichthyology ECI X„ 5. – 10. September 2004, Thallin.

· Freyhof, J. & Kottelat, M.: Most threatened freshwater fishes of Europe: An overview. (oral)

· Freyhof, J.: National responsibility, red lists and IUCN categories: The German approach. (oral)

· Nolte, A.; Stemshorn, K., Freyhof, J. & Tautz, D.: Hybrid zones in Cottus and the genetic architecture of divergence. (oral)

· Naseka, A. M. Spodareva, V. V., Bogutskaya, N. G. & Freyhof, J.: A comparative faunistic study of the Kuban River fishes: Historical reasons of species composition and zoogeographical originality. (oral)

· Slechtova, V., Bohlen, J., Freyhof, J., Persat, H. & Delmastro, G.: Origin and dispersal of Adriatic Cottus: A peri-alpine clade indicates geneflow between major drainage areas. (oral)

· Slechtova, V., Bohlen, J. & Freyhof, J.: Extracting clades from the "Loach Bush": Botiidae (Teleostei: Cobitoidea). (oral)

„9. International Symposium on the Biology and management of Coregonid Fisches” 21. - 27. August 2005, Olsztyn, Poland.

· Freyhof, J. & Schöter, C.: The houting, Coregonus oxyrinchus (L.), a globally extinct species from the North Sea basin (Salmoniformes: Coregonidae). (oral)

· Freyhof, J. & Mehner, T.: Trophic morphology of sympatric Baltic ciscoes (Coregonus albula species group). (oral)

· Palm Helland, I., Freyhof, J. & Mehner, T.: Depth distribution of sympatric pairs of ciscoes (Coregonus spp.). (poster)

„5. Tagung der Gesellschaft für Ichthyologie (GfI)„ 22.-24. September 2005, Bonn/Germany.

· Freyhof, J.: Endemische Fische der Alpenseen – Schatztruhe oder Stinkkiste? (oral)

· Freyhof, J.: Rote Liste Süsswasserfische 2008: Vorstellung des Kriteriensystems. (oral)

· Özulug, M. & Freyhof, J.: Revision der Mairenken der Alburnus chalcoides-Gruppe (Cyprinidae) unter besonderer Beachtung der Populationen der Westürkei. (poster)

· Stelbrink, B. & Freyhof, J.: Reduktion von Schuppen und Kopfkanälen bei Pomatoschistus canestrini (Ninni, 1883) (Gobiidae). (poster)

“Hybridization in Animals” 12-15.October 2006, Frankfurt, Germany

Freyhof, J., Lieckfeldt, D., Pitra, C. & Ludwig, A.: Distant, intergeneric hybridization in Dalmatian cyprinids (Teleostei: Cyprinidae). (oral)

“3rd International Conference: Loaches of the Genus Cobitis and Related Genera, Biology, Systematics, Genetics, Distribution, Ecology, Conservation” 24-29. September 2006, Sibenik, Croatia

Freyhof, J., Stelbrink, B.: Red List of European Cobitidae: Listing of most threatened species for conservation priority. (oral)

Freyhof, J., Stelbrink, B., Özulug, M. & Economidis. P. S.: The first record of Cobitis puncticulata from Europe with comments on its conservation status (Teleostei: Cobitidae). (poster)

Something interesting? Questions? Just sent me a mail.

to head of document